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Scattered across the pristine isolation of Nevada are many
productive Ordovician-age fossil localities roughly 490 to 440
million years old, and two of the more significant sites can
be visited in the Toquima Range. At what many fossil seekers
call Ordovician Canyon, for example, paleontology enthusiasts
can find a plethora of well preserved invertebrate animal remains
from the Middle Ordovician Antelope Valley Limestone, including
silicified brachiopods, bryozoans, sponges, cystoid echinoderms,conodonts,
trilobites, gastropods, pelecypods, cephalopods and ostracodes.
A second site paleontology seekers refer to as Graptolite
Summit gives collectors a chance to find a fossil type many hobbyists
rarely see outside of a textbook or popular guide to paleontology--the
intricately designed graptolite, a colonial organism most often
referred to as an extinct variety of hemichordate (or primitive
chordate). The specimens at Graptolite Summit occur in a rock
deposit called the Vinini Formation, which has been dated by
geologists as Early to Late Ordovician. The Vinini is an incredibly
widespread unit throughout Nevada, a dominantly siliceous assemblage
of shales, siltstones, cherts and quartzites that bear sporadic
occurrences of abundant graptolites. With the possible exception
of the type locality (where a geologic rock formation was first
named and described in the scientific literature), this particular
area surrounding Graptolite Summit is probably the most intensively
investigated graptolite-yielding section in all of the Vinini
Formation. Over the decades it has provided professional paleontologists
and amateurs alike with myriads of identifiable graptolites,
in addition to common inarticulate brachiopods and carapaces
belonging to a peculiar species of extinct cructacean called
Caryocaris, whose oval-to D-shaped exoskeletons up to
an inch across appear to be confined throughout the world to
shales in which graptolites are the dominant fossil specimens
preserved.
| Click on
the images for larger views. Here are two fossil-types collected
from the Toquima Range, Nevada: brachiopods and a graptolite.
From left to right: left--a silicified pedicle valve from the
brachiopod Hesperorthis affin. H. matulina Cooper
(resting on its natural limestone matrix), 10mm acrosss, from
the Middle Ordovician Antelope Valley Limestone, Ordovician Canyon,
Nevada; middle--an oval graptolite colony, 17mm long, called
Phyllograptus anna from the Lower to Upper Ordovician
Vinini Formation near Graptolite Summit, Nevada; right--a silicified
pedicle valve from the brachiopod Plectorthis cf. P.
perplexus (Ross), 10mm across, (still on its natural limestone
matrix) from the Middle Ordovician Antelope Valley Limestone,
Nevada. |
Both fossil areas are easily and safely reached. Still,
collectors must not be lulled into dangerous complacency. The
important thing to remember is that this part of Nevada remains
one of the most remote sectors in all the Great Basin. Should
a genuine emergency emerge while in the deep backcountry, medical
and mechanical assistance will certainly be a long time in arriving,
even if your situation has been relayed to the authorities over
Citizens Band Radio. For this reason, it is recommended that
visitors travel to the fossiliferous regions in the Toquima Range
only in a reliable four-wheel drive vehicle, obeying all of the
necessary rules that apply to back country travel; carry plenty
of water, emergency provisions, cold-weather clothing, spare
fan belts, and medical supplies. And by all means notify the
authorities in the nearest community of your whereabouts, remembering
to check back in with them upon leaving the area.
The Graptolite Summit locality rests directly atop the
Lower to Upper Ordovician Vinini Formation, which is locally
loaded with all kinds of interesting graptolite remaims. Here
in the Toquima Range the Vinini Formation has been measured by
geologists at some 6,000 feet thick. It is predominantly a siliceous
accumulation of thin-bedded black chert, quartzite, red to black
siltstone and shale, dark limestone, and even some minor interbeds
of pillow lavas, presumably formed on the ancient Ordovician
ocean floor when hot magmatic extrusions came in contact with
obviously much colder marine waters; identical kinds of lavas
develop today in ocean waters near sites of sea-floor-spreading,
where the so-called Mid-Atlantic Ridge produces new Earth crust
through the upwelling of superheated magmas.
| Click on the
image below for a larger view. Looking northwestward from the
fossil graptolite locality in the Middle to Upper Ordovician
Vinini Formation near Graptolite Summit, Nevada. The pastel-colored
to black shaley siltstones which sandwich a prominent bed of
whitish-brown quartzite (a knob of the quartzite is visible at
the extreme upper right) near Graptolite Summit furnish paleontology
enthusiasts with loads of fascinating graptolite remains, an
extinct variety of hemichordate that reached its zenith of adaptation
in the Ordovician Period. |
Most
of the graptolites occur in pastel-colored shaley siltstones
of the Vinini Formation. Numerous scientific crews have worked
these fossiliferous siltstones in the vicinity of Graptolite
Summit over the decades, periodically entrenching the thin-bedded,
poorly exposed sedimentary layers to a depth of several feet
in search of productive graptolite layers. Depending on the degree
of erosion inflicted by wintertime's snow drifts in the Toquima
Range, remnants of their abandoned excavations may be visible
just up slope from a prominent bed of whitish-brown quartzite
interbedded in the section. The quartzite layer is called a key
marker bed by stratigraphers, because it conveniently separates
the lower member of the Vinini Formation from the upper member.
All of the rich graptolite horizons in the vicinity of Graptolite
Summit occur within a rather restricted interval of shaley siltstones
and shales some 400 to 600 feet thick, a productive section that
happens to straddle that massive, distinctive bed of quartzite.
In general, rocks to the southeast of the quartzite marker bed
are younger than those to the northwest, but the fact remains
that most of the shales and siltstones that sandwich the quartzite
horizon yield rare to abundant graptolite specimens.
The most efficient way to find graptolites here is to remove
sizable chunks of the varicolored to black shaley siltstones,
then carefully split the rocks along their natural bedding planes
(remember to wear protective eye gear). In doing this, most collectors
soon realize the despite such a prominent presence of graptolites
in the rocks, the fossils are sometimes difficult to spot on
the fine-grained matrix, a number of them appearing as small
silvery sheens on the surface of the shales. Do not become discouraged.
What you have discovered is what graptolite specialists have
known for ages--that the vast majority of specimens project to
the unaided eye what has become known as a "traditional
graptolitic aspect of preservation." That silvery sheen
found glinting out at you when the sunlight strikes the surface
of the rocks at just the right angle represents a 475-million-year-old
graptolite colony whose original skeleton has been compressed
through geologic time. Most specimens range anywhere from a quarter
to an inch and a half in length and, depending of course on the
particular genera of graptolites unearthed, can present a fascinating
variety of distinctive shapes and sizes to study. Phyllograptus
graptolites, for example, one of the more obvious types found
near Petes Summit, grew, oval, roughly football-shaped colonies
a little over a half inch long. Also present are the blade-like
Orthograptus and Climacograptus, plus wishbone-shaped
Didymograptus and slingshot-like Dicranograptus. Other
genera available in the Graptolite Summit rocks include Clonograptus,
Tetragraptus, Isograptus, Glyptograptus, Dicellograptus, Paraglossograptus,
Pterograptus, Amplexograptus, Durangograptus, Callograptus and
Cardiograptus.
In addition to the graptolite remains in the Vinini Formation
near Graptolite Summit, two other fossil types can be encountered
in the shales and siltstones--inarticulate brachiopods and Caryocaris
crustaceans. Such remains are far less abundant than the
graptolites, though. Collectors interested in finding them would
be advised to explore as many of the shale deposits as possible,
splitting heaps of the easily separated layers wherever you go.
And don't be shy about exploring the little gullies and ravines
in the Graptolite Summit district--many graptolites, for example,
can be found in the poorly exposed shales and siltstones that
seem to hide in the most improbable-appearing areas.
| Click on
the images for larger views. Here are two varieties of graptolites
from the Middle Ordovician Vinini Formation exposed near Graptolite
Summit, Nevada. At left is a dendroid-type graptolite, 15mm long,
called Callograptus sp.; at right is the slingshot-like
graptolite called Dicranograptus spinifer Elles and Wood,
22mm long, which has been naturally preserved with an unusual
reddish coloration, presumably through replacement by the mineral
limonite. |
Graptolites first appear in the geologic record during
the middle stages of the Cambrian Period, some 505 million years
ago. Even though they persisted all the way up to the late Mississippian
age, or roughly 325 million years ago, most species of graptolites
had already become extinct by the latest Devonian Period 35 million
years earlier. Graptolites achieved their highest degree of success
during the Ordovician Period, when they attained worldwide distribution
by adapting with ingenuity to three distinct modes of life. One
order of graptolite, for example--the fan to leaf-shaped dendroids--led
a sessile life attached to the sea floor, apparently straining
the marine waters for microscopic organisms. Another type developed
a special flotation device which allowed the graptolite colony,
termed a rhabdosome, to drift in the open ocean; and a third
kind solved its own planktonic challenge by attaching itself
to floating strands of seaweed to hitch a free ride through the
open ocean in search of better feeding grounds; presumably it
too strained the sea waters for microscopic particles of food.
In all three examples of graptolitic adaption, the actual
colonial animal lived inside the minute rows of cups called thecae
that developed along each individual segment of the rhabdosome;
technically, these segments are called a stipe. The tiny saw-tooth
compartments that housed the graptolite animals along the stipe
show to best advantage under magnifications of ten or more power.
Thus, a good-quality hand lens is indispensable in order to gain
a detailed and aesthetic appreciation of your finds.
The exact zoological classification of graptolites has
presented a serious challenge to paleontologists. Early investigators
referred graptolites to such disparate groups as coelenterates
or bryozoans; yet, there certainly was no unanimity of opinion
among fossil specialists throughout the 19th century. The breakthrough
came when some perfect, three dimensional specimens were etched
out of cherts using powerful brews of acids around 1948. Paleontologists
then realized that the graptolite colony most closely resembled
the modern pterobranch, a tiny marine hemichordate, which by
definition is a primitive chordate whose notochord (a spine-like
notch) is restricted to the basal part of the head.
That explanation seemed to satisfy most paleontologists.
Even the basic idea that the graptolite was an extinct colonial
organism went completely unchallenged until 1989 when Noel Dilly,
a marine biologist in London, suggested that the graptolite had
not died out, that a single species had survived the Paleozoic
Era and was alive and well on Earth today.
What Dilly had identified was a dime-sized colony dredged
up by a French team from 800 feet off of New Caledonia in the
South Pacific. Dilly called it Cephalodiscus graptiloides--the
sole surviving member of the graptolitic race, he claimed. Dilly,
who published his ideas in the Journal of Zoology a number
of years ago, also reported that while on vacation he actually
witnessed his "living graptolites" cavorting in the
warm, shallow waters off the coast of Bermuda. He speculates
that his living fossils are "survivors of the main group
who hung on in places where there hasn't been massive change
in the environment in over 300 million years."
The suggestion is a novelty, at best. By Dilly's own admission
the chemical structure of the graptolite rhabdosome and that
of his living fossil is only "similar," not identical.
What generated most of the early enthusiasm for the theory was
that his Cephalodiscus apparently possesses an extended
spine-like protrusion from the main colony, a structure similar
to what paleontologists call a nema on fossil graptolites. Modern
pterobranchs, with which the graptolite is most often compared,
do not develop such a needle-like projection, or nema, so the
recent identification of a colonial hemichordate that does seem
to bear a nema created quite a short-lived stir among paleontologists.
One of the main problems with the entire concept is that Dilly's
Cephalodiscus graptiloides is not the only species in
its genus, and it's the only one to produce a nema--a structure
which may not be directly analogous to the structures graptolites
developed.
| Click on
the images for larger views. Here are two varieties of graptolites
from the Middle Ordovician Vinini Formation near Graptolite Summit,
Nevada: at left is a Climacograptus sp. graptolite, 33mm
long; at right are several thread-like Clonograptus flexilis
(J. Hall) graptolites, the longest of which is 20mm. |
After collecting graptolites near Graptolite Summit, visitors
will want to visit the spectacular fossil exposures of the Antelope
Valley Limestone at Ordovician Canyon in the Toquima Range. Here,
the Middle Ordovician Antelope Valley Limestone is roughly 950
feet thick, yielding prodigious numbers of fossilized shelly
creatures. The productive limestone layers near the mouth of
the canyon (referred to as the Mill Canyon Sequence by geologists;
two miles from the mouth, geologists call the strata the June
Canyon Sequence) consist principally of silty to finely crystalline
limestones that weather into shades of dark gray, medium gray,
grayish orange, grayish yellow, yellow gray, brownish orange
and yellowish orange. The most fossiliferous exposures occur
northeast of the mouth of Ordovician Canyon, but productive horizons
can be discovered through the canyon corridor up to two to two
and a half miles west of the mouth.
| Click on
the images for larger views. At left is the view looking westward
from the mouth of Ordovician Canyon, Nevada, to the distinctive,
bold outcrops of the fossil-bearing Middle Ordovician Antelope
Valley Limestone; at right, a collector (faintly visible at middle
right of image) prospects the talus slopes just north of the
mouth of Ordovician Canyon for Middle Ordovician invertebrate
fossils in the Antelope Valley Limestone. |
| Click on
the image below for a larger view. Here is a slab of shaley limestone
from the Middle Ordovician Antelope Valley Limestone, Ordovician
Canyon, Nevada, which bears several silicified pedicle valves
of the brachiopod Plectorthis cf. P. perplexus (Ross);
all of the specimens are roughly 10mm across. |
Many
collectors like to concentrate their attention along the moderate
talus slopes immediately north of the mouth. Here can be found
infrequent to relatively common brachiopods and gastropods, in
addition to abundant cystoid echinoderm debris, or small crinoid-like
ossicles whose precise identification is impossible owing to
the fragmentary nature of the material. Since these easily accessible
exposures have been probed by eager collectors for decades, the
richest limestone layers, those yielding the greatest diversity
and abundance of specimens, can now be found only in the rugged
terrain farther north of the road. In this area the Antelope
Valley Limestone is more reliably fossiliferous, yielding a genuinely
remarkable assemblage of nicely preserved remains--all of them
thoroughly silicified, by the way, replaced by silicon dioxide.
Such a style of preservation means that collectors can immerse
the fossiliferous calcium carbonate matrix in a diluted acid
batch, dissolving away the limestones to leave intact, perfect
specimens in the residues.
| Click on
the image below for a larger view. The prominent outcrops of
the Middle Ordovician Antelope Valley Limestone immediately north
of the mouth of Ordovician Canyon, Nevada, yield common to abundant
silicified invertebrate fossils, including brachiopods (arguably
the most common fossil type encountered), echinoderm debris,
trilobites, sponges, ostracodes, conodonts, bryozoans, pelecypods,
cephalopods and gastropods. |
In addition to brachiopods, gastropods and cystoid echinoderms,
other specimens identified from the Antelope Valley Limestone
include ostracodes, trilobites, conodonts (acetic acid must be
used to dissolve out the phosphatic conodont elements), cephalopods,
sponges, pelecypods and bryozoans. Most of the fossil groups
appear to occur within distinct and separate zones within the
Antelope Valley Limestone. Some of the protruding limestone ledges,
for example, yield many ostracodes, while others bear plentiful
trilobite fragments, brachiopods, sponges, gastropods, or cephalopods.
Conodonts, on the other hand, may show up in the residues of
limestones collected throughout the entire thickness of the formation,
appearing as minute (only one to three millimeters in length,
or less than an eighth of an inch) tooth-like specimens that
originally served as a unique feeding apparatus of an early,
primitive eel-like chordate.
The Toquima Range localities offer collectors a superlative
selection of well preserved Middle Ordovician fossil specimens
some 475 million years old. Along with several specific localities
in Utah, Ordovician Canyon may well be one of the most fossiliferous
Ordovician sections in all the Great Basin. Add to that the profusion
of fascinating graptolites near Graptolite Summit and you have
an extensive fossil field that begs to be explored--preferably
during mid Spring through early Fall when the weather conditions
most reliably favor a comfortable experience.
Both Toquima Range fossil localities lie within a designated
United States national forest. This means that they are administered
by the United States Forest Service, not the Bureau of Land Management,
even though the sites occur on public lands. In the past, hobby
fossil collecting has been allowed to go on here without the
need of a special use permit. Just to be on the safe side, though,
you might want to contact the local Forest Service office before
any visit is made to the Toquima Range.
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